Motif Tools
Family atlas / C2H2 zinc finger

KRAB zinc-finger proteins (KZFPs)

C2H2 zinc finger587 motifsaka KZFP, KRAB-ZFP, KRAB-ZNF
TACGCTCCCGATTCAGAGTC
representative: M07723_2.00 · data consensus GCTCCCTGC
members not one motif · 0.37no single consensus61 in vitro224 ChIP-only14 references, all PMIDs verified

KRAB zinc-finger proteins (KZFPs) are the largest family of DNA-binding proteins in the human genome: roughly 350 to 380 genes, most named only ZNF followed by a number. Each combines a KRAB repression domain, which recruits the corepressor TRIM28/KAP1 and the histone methyltransferase SETDB1, with a tandem array of C2H2 zinc fingers that reads a long, protein-specific stretch of DNA. Most of those stretches lie inside transposable elements, and the family has expanded and diversified alongside successive waves of element invasion. This atlas entry is therefore a bucket, not a motif: its 587 matrices come from 304 different proteins, run from 4 to 38 bp, and mostly do not resemble one another. A hit identifies one protein's site, not a family motif.

The factors

Human KZFPs number about 350 in the original survey and 378 in the most complete census, the largest group of transcription factors encoded by tetrapods (Imbeault et al. 2017; de Tribolet-Hardy et al. 2023) 28273063,37730438. The architecture is uniform: an N-terminal KRAB domain, a variable linker, and a tandem array of 4 to more than 30 C2H2 fingers. The KRAB domain binds the corepressor KAP1/TRIM28 (Friedman et al. 1996) 8769649, which in turn scaffolds the H3K9-specific methyltransferase SETDB1, HP1 proteins and DNA methyltransferases, producing H3K9me3 heterochromatin over the bound site (Schultz et al. 2002) 11959841. The genes sit in clusters, heavily on chromosome 19, built by repeated gene and segment duplication; the family traces back more than 400 million years, to a common ancestor of coelacanths and tetrapods (Imbeault et al. 2017) 28273063. Selection has acted specifically on the finger residues that touch DNA, so recent paralogues diverge in specificity while their scaffolds stay similar (Emerson and Thomas 2009) 19119423. Not all are simple repressors: across 101 KZFP interactomes, those conserved up to placental mammals essentially recruit KAP1, while a subset of more ancient ones interact instead with genome-architecture and RNA-processing factors (Helleboid et al. 2019) 31403225. Expression is not restricted to stem cells. This entry carries 304 distinct gene symbols, 191 of which have two or more matrices.

The motif

There is no family consensus and one should not be invented. Every protein's finger array differs, and each finger specifies roughly three base pairs through a few residues in its recognition helix, so a 10-finger protein can specify 30 bp. That is what the data show: median consensus length 17 bp, 360 of 587 matrices at least 15 bp, 214 at least 20 bp, median information content 16.3 bits, and 149 matrices above 20 bits. The longest is a 31 bp, 50.7-bit model for ZNF550. Family cohesion is 0.37, against values above 0.8 for entries that really are one motif, and the listed representative (CIS-BP M07723, GCTCCCTGC) is just the matrix nearest the centre of a diffuse cloud. Typical examples: ZNF558 ATTTGATGAAAGACATAAATCTACAAATC (29 bp), ZNF611 TCTGGGATGTGAGGAGCGCCTCTGC (42 bits), ZNF75D TGTAATGAATGTGGGAAAGCCTTCA. Recent paralogue pairs are not separable: ZNF417 and ZNF587 share TGGCGCCCAACATGGG, and the same holds for ZNF75A/ZNF75D, ZNF354A/ZNF354B and the ZNF33A/ZNF33B archetypes. A minority of matrices have the opposite problem, being short or low-complexity and therefore non-specific: 24 are under 8 bp, and models for ZNF212 (a TC repeat), ZNF613 (poly-T) and ZNF746 (poly-C) carry high nominal information but no factor-specific content.

Where it acts

Overwhelmingly inside transposable elements: endogenous retroviral LTRs, L1, SVA and Alu. Most KRAB-containing C2H2 proteins bind specific endogenous retroelements, from currently active families to ancient ones, and also show widespread binding at regulatory regions (Najafabadi et al. 2015) 25690854. Critically, most element targets have lost all transposition potential, so KZFPs are using conserved element fragments as regulatory platforms long after any arms race ended, creating a largely species-restricted layer of epigenetic regulation (Imbeault et al. 2017) 28273063. The chromatin context is usually H3K9me3-marked and closed: many KZFPs bind within closed chromatin, each recognizing a distinct set of repeats (Razavi et al. 2026) 42557260, so these sites are routinely absent from accessibility-based catalogues. A separate niche is the 3' end of the zinc-finger genes themselves, where ZNF274 recruits KAP1 and SETDB1 (Frietze et al. 2010) 21170338. Occurrence counts track the copy number of the target element subfamily, from hundreds to tens of thousands of loci.

Biology and disease

Depleting TRIM28 in human or mouse embryonic stem cells derepresses a broad range of transposable elements, which is the baseline phenotype for the family (Imbeault et al. 2017) 28273063. Individual proteins show how element control becomes gene control. ZNF558 evolved as an L1 suppressor but now regulates a single target, the mitophagy gene SPATA18, in human but not chimpanzee forebrain progenitors, with its own expression set by a variable tandem repeat that differs between the two species (Johansson et al. 2022) 34624206. More broadly, KZFPs act as context-specific switches that gate element-derived regulatory modules, and mutations in KZFP genes are an emerging cause of human developmental disorders (Davis et al. 2026) 41720928; Bruno et al. 2019 review the arms race and its consequences for mammalian traits 31518518. Imprinting (ZFP57), pancreatic development (ZNF808) and the SVA/L1 restriction factors ZNF91 and ZNF93 have separate entries.

Reading a hit

Start from the gene symbol on the matrix, not the family. Then check the evidence: of the 285 HOCOMOCO models here, 224 are ChIP-only (evidence code P) and 160 are graded C, with only 49 at grade A, and in vitro codes (S, M) are rare. A ChIP-derived KZFP motif is in practice the consensus of the element the protein sits on, so a match may mean only "this is that element". Long, high-information motifs will not arise by chance in a small interval, but they will match every copy of the parent element, so genome-wide enrichment is usually a statement about repeat content. Seventeen matrices here have their closest neighbour in a different family: three resemble ETS motifs, two each MAZ, CTCF, ZNF24 and TBX, and one each STAT, YY1, HNF4, E2F, ZNF384 and ZFP57. That is the expected failure mode of ChIP-derived models, and a reason to check short KZFP hits against the better-characterized families. Finally, most KZFPs are expressed at low levels in restricted contexts, so a sequence match without expression evidence in your cell type is weak.

How this was worked out

  1. 1991mediumsequence comparison, zoo blots, northern blots2023909

    A conserved 75-residue N-terminal region, named the Krüppel-associated box, was found in about a third of the hundreds of human Krüppel-like zinc-finger genes, defining the family as a subfamily rather than a scatter of individual genes.

    Still open then: Nothing was known about what any of these proteins bound, or how the KRAB domain repressed.

  2. 1996mediumtwo-hybrid cloning, repression assays8769649

    KAP-1/TRIM28 was cloned as the corepressor that binds the KRAB domain and is required for its repression, giving the whole family a single shared effector.

    Still open then: How KAP-1 silenced chromatin, and which genomic sites KRAB proteins occupied, were both unknown.

  3. 2002highbiochemistry, in vitro methyltransferase assays, immunofluorescence, ChIP11959841

    KAP-1 was shown to bind SETDB1, an H3K9-specific methyltransferase, and KAP-1, SETDB1, H3K9me3 and HP1 were found together at a silenced promoter, supplying the chromatin mechanism behind KRAB repression.

  4. 2014highcomparative genomics, ancestral-sequence reconstruction, reporter and repression assays25274305

    ZNF91 and ZNF93 were shown to have evolved to repress SVA and L1 elements shortly after those elements began to spread, and the L1PA3 subfamily was shown to have escaped ZNF93 by deleting its binding site, establishing the arms-race model for the family's expansion.

    Still open then: Whether the arms race described two primate genes or the whole family was untested, as targets were known for only a handful of KZFPs.

  5. 2015highbacterial one-hybrid, protein-binding microarray, ChIP-seq, inferred recognition code25690854

    A recognition code derived from thousands of C2H2 finger domains, combined with ChIP, gave motifs for large numbers of zinc-finger proteins and showed that most KRAB-containing members bind specific endogenous retroelements while also binding regulatory regions.

    Still open then: Motifs inferred from a code or from ChIP peaks could not separate the protein's own specificity from the sequence of the element it sits on.

  6. 2017highChIP-exo in human cells, phylogenetics28273063

    ChIP-exo across a large fraction of human KZFPs put the root of the family at a common ancestor of coelacanths and tetrapods and showed that most bound transposable elements have lost all transposition potential, revising the arms race from an ongoing conflict into a largely finished one whose products are reused as regulatory platforms.

    Still open then: It remained unclear how many bound element copies are actually repressed in any given cell type.

  7. 2019mediumaffinity purification-mass spectrometry31403225

    Interactomes for 101 KZFPs showed that members conserved up to placental mammals essentially recruit KAP1, while a subset of more ancient ones instead bind genome-architecture and RNA-processing factors, so the family is not uniformly a set of heterochromatin repressors.

  8. 2026mediumChIP-seq of tagged factors in living cells; review of human genetics42557260 41720928

    Systematic profiling of poorly characterized human transcription factors found that many KZFPs bind within closed chromatin at protein-specific sets of repeats, sites that accessibility-based catalogues miss, and KZFP mutations were argued to be an emerging cause of human developmental disorders.

    Still open then: For most individual KZFPs there is still no loss-of-function phenotype, so the functional consequence of this binding is unmeasured.

Still unresolved

How much to trust a hit

This entry is a family of proteins, not a motif: cohesion 0.37, with 587 matrices from 304 genes whose sites are largely unrelated. Never report it as a consensus. Most models are ChIP-derived (224 of 285 HOCOMOCO entries, 160 graded C), and for KZFPs that usually means the matrix is the consensus of a transposable element, so a hit may amount to repeat annotation rather than evidence of binding. Recent paralogue pairs such as ZNF417/ZNF587 share a motif outright. Seventeen matrices most closely resemble motifs of other families, including ETS, CTCF and MAZ. Check the gene symbol, the evidence code, the repeat track, and whether the protein is expressed in your cells.

References

587 motifs in this family — JASPAR 51 · HOCOMOCO 285 · CIS-BP 240 · Vierstra 11
motifdatabaseconsensusbits
ZN550.H14CORE.0.P.CPCHOCOMOCOCTTCTGCACTGCCCTAGCAGAGGTTCTCCAT50.7
M07609_2.00CIS-BPATTgCGCCACTGCACTCCAGCCTGGGCaAC49.9
ZN441.H14CORE.1.P.CPCHOCOMOCOTTGAGACGGAGTCTCGCTCTGTCGCCCAGGC45.9
M07768_2.00CIS-BPGAGTGGCTTTTTCTACCTCACAATACAACC45.3
ZN649.H14CORE.0.P.CPCHOCOMOCOGATATAATCTCCTGGTGTGCCGTTT45.2
M07625_2.00CIS-BPTCTCTCTCTCTCTCTCTCTCTCTCTCTCTC44.4
ZN611.H14CORE.1.P.CPCHOCOMOCOTCTGGGAtGTGAGGAGCGCCTCTGC42.4
ZNF90.H14CORE.0.P.CPCHOCOMOCOAcGCCTCTCTGATTATTCACCCA40.7
M07742_2.00CIS-BPACTCTTTCTCTGCTGCAACACCTGCTGTT39.8
ZN443.H14CORE.0.P.CPCHOCOMOCOTAGCGGGTGTTTTTCCTTGACAC39.4
ZN675.H14CORE.0.P.CPCHOCOMOCOgGAGGCCTAGGAGGAAAAAATGGTT39.3
ZN778.H14CORE.1.P.BPBHOCOMOCOcgGCATCTGGTGAGGGCCTTCTTGCTGtgTC38.8
ZN184.H14CORE.0.P.BPBHOCOMOCOCCTCtCCAgCACtTGTTgTTtcCtGaCTTTT38.7
M07643_2.00CIS-BPTTTCTAAATGCAGCAACCTbTgTTcTTCA38.4
ZN430.H14CORE.0.P.CPCHOCOMOCOCACaTCTTACATGGtGGCAGGCA37.5
M07640_2.00CIS-BPCTCTGACCCCgCCTCCctTCTAAA37.5
Z585B.H14CORE.0.P.CPCHOCOMOCOGAGGCCTAGGAGGAAAAAATGGTT37.4
ZN562.H14CORE.0.P.CPCHOCOMOCOTTTTCTCAGCAAGGCAATTTTacTT37.4
ZN254.H14CORE.0.P.CPCHOCOMOCOAAGATGTAGGCTGGGAGGCTA37
ZN586.H14CORE.0.P.CPCHOCOMOCOGAGGCCTAGGAGGAAAAAATGGT36.5
ZN564.H14CORE.0.P.CPCHOCOMOCOTTGGACTTCCCAGCCTCCAGAAC36.4
ZN578.H14CORE.0.P.BPBHOCOMOCOCCTGAGGcGACATACATCCTC36.4
Z705E.H14CORE.0.P.BPBHOCOMOCOGGGAGAGGGAGAGGGAGAGGG35.4
M07564_2.00CIS-BPGCGTtCtCAGCCCaAGGCATGCCAGAGG35.3
ZN627.H14CORE.0.P.CPCHOCOMOCOGTAGGTCTCAACAGTGGGCTTAA35.1
M07660_2.00CIS-BPCCGcGCCTAcCTGaGGGCGCgGGCcTGGgG35.1
ZN534.H14CORE.1.P.CPCHOCOMOCOAgAAgGGGttggGGGGTtCTTGCCCCcCacc34.9
ZN248.H14CORE.0.P.BPBHOCOMOCOATaCCATGTTCATGGATtGGAAG34.6
M07682_2.00CIS-BPCACACccTCAcCAtaaGAGctGgCGCCCGC34.5
ZN302.H14CORE.0.P.BPBHOCOMOCOGGAACCACAGTCACTCAACTACA34.2
ZN736.H14CORE.0.P.CPCHOCOMOCOGCCCATAAAAaCCCCAGACTCAG34
M07670_2.00CIS-BPCcacTTTTATACCCAaCCCtaaTatCCa33.8
ZN75D.H14CORE.1.P.BPBHOCOMOCOTGTAatGAATGTGGGAAAGCCTTCA33.7
ZN112.H14CORE.0.P.CPCHOCOMOCOTGCCAATTtCtgTGGTGTAAATAC33.4
ZN100.H14CORE.0.P.BPBHOCOMOCOGCACTTCTTACATGGtGGcGGCA32.9
ZN565.H14CORE.0.P.CPCHOCOMOCOaGCcGCCATGtTGTGAGGAAGCCCAAGCagC32.9
M07708_2.00CIS-BPTTTTGCTGGaTATAGtATTCTTGGTT32.8
ZN37A.H14CORE.0.P.BPBHOCOMOCOGbGAGGGAGCTGGGGTATTTATA32.5
ZNF558 (MA2335.1)JASPARAtTTGATGAAAGACATrAATcTACAvATc32.3
ZNF92.H14CORE.0.P.BPBHOCOMOCOATGTCCAGGCAGAAGTtTGCT32.2
ZN529.H14CORE.0.P.BPBHOCOMOCOTcTttACtCTTtATCCAGcTTcCCccacTG32
ZN805.H14CORE.0.P.CPCHOCOMOCOGATGsGGCCACAAGCCAAGGAATGC31.9
M07777_2.00CIS-BPGCCAATTtCcgTGGTGTAAAT31.9
M07720_2.00CIS-BPTGTGTGTGtGggcaTGTGTgTGtgcgaGT31.6
M07782_2.00CIS-BPCaAAaAAAAAaAaAAtAAhAAgCAaAcaA31.6
ZNF10.H14CORE.0.P.CPCHOCOMOCOGAcCCCTACCTCACACCATATAC31.4
M07766_2.00CIS-BPGcTcTatTgTTtGGTGCATaTATgTTTAttATTgTTAT31.1
ZN436.H14CORE.0.P.CPCHOCOMOCOGGAaGGCTTCCTGGAGGAGGtGg30.7
M07675_2.00CIS-BPAACAAcaCTGCAaTGAACATC30.6
ZNF74.H14CORE.0.P.CPCHOCOMOCOAAAcAAtaCTGcAATGAACATC30
M07779_2.00CIS-BPCCntCCcCCCTCTtCtyCCgGCCTGTGGCC29.8
ZNF81.H14CORE.0.P.CPCHOCOMOCOgGTACAACCACTaTGGAAA29.5
AC0381:ZNF33A:C2H2_ZFVierstraTCTCGGCACTTTGGGAGGCC29.2
ZN490.H14CORE.0.P.CPCHOCOMOCOTGaAGgCAGCAtAtcaTTGGGTC29.1
ZN250.H14CORE.0.P.CPCHOCOMOCOCAGTACCATgCTGTTTTGaTTAC28.8
ZNF8.H14CORE.1.P.BPBHOCOMOCOTGTGGTATATCCATACAATGGA28.8
M07749_2.00CIS-BPTTTAAGCCCACTGTTGAG28.8
ZN304.H14CORE.0.P.CPCHOCOMOCOCCwtCCCCAGTcgAGCCTTCAGATGAgacCa28.2
ZN560.H14CORE.0.P.CPCHOCOMOCOCCACTGCACTCCAGCCTGGGcgA28.2
M08234_2.00CIS-BPGTCAACTTGACTGGGCCA27.8
M07776_2.00CIS-BPTTACCatTTaaATTtCTTTcTTTtTtTT27.6
ZFP90.H14CORE.1.P.CPCHOCOMOCOcCCTCTtAgCACTGCTTTAGCTG27.5
M07740_2.00CIS-BPTcCTgATTccacaCTcAAAhaTaAcTTT27.5
ZN233.H14CORE.0.PG.APGAHOCOMOCOATAGCcaAACAAAAGGCAGCAGAAA27.4
ZN136.H14CORE.0.P.BPBHOCOMOCOTTTGCTGGaTATAGaATTCTTGG27.2
M04575_2.00CIS-BPTTTCctGTCTTTTTTCT27.2
ZN135.H14CORE.0.P.CPCHOCOMOCOAGcCTCGACCTCCTGaGCTCAAG26.7
M07648_2.00CIS-BPgCTTnCaAnAAaaATTTAACTCCCabyTC26.6
ZN791.H14CORE.0.P.CPCHOCOMOCOccttTgTATgTaaTtAgTTAtTT26.4
ZN433.H14CORE.1.P.CPCHOCOMOCOTtgCtggattACtAcTGtgGTCaTttGGaGg26.3
M07568_2.00CIS-BPgGGGGcCCtchngaATGGGATTAGTGCCCT26.3
ZNF382 (MA1594.1)JASPARAGGgGAcATCACtACaGAtCCCAC26.2
ZN284.H14CORE.0.P.CPCHOCOMOCOGTGCAGTGGTGCGATC25.6
ZN154.H14CORE.0.P.CPCHOCOMOCOGGAAGACCTCCtgGACaAAGAC25.5
ZN772.H14CORE.0.PG.APGAHOCOMOCOaGTCAACTGATTAGgrACcTT25.4
ZN558.H14CORE.0.P.BPBHOCOMOCOTGATGAAAGACATrAATCtACAgAT25.3
ZNF8 (MA1718.1)JASPARGTATGGATATACCACAtTTT25.2
ZN584.H14CORE.0.P.BPBHOCOMOCOATTTCAaAATTGCTATGGaCCA25.2
Z354A.H14CORE.0.P.BPBHOCOMOCOTTTAgtCCATTTACATTTAATGT25.1
M07769_2.00CIS-BPAGCCATCCtTtGCTGvCaGgg24.9
M07590_2.00CIS-BPTtTkTcTTttattTtTtTTtTTTTTTTTTTT24.8
M07737_2.00CIS-BPGvTGAGCTCTTAAGGCCAC24.6
ZN875.H14CORE.0.P.CPCHOCOMOCOGATAtgaTGCACTgaGgAGGACAcA24.4
ZN331.H14CORE.0.P.BPBHOCOMOCOGGCtGGGCTCAGCTGGGtGGTTC24.3
M07700_2.00CIS-BPATGtTGTGAGGAAGCCCA24.3
M08292_2.00CIS-BPTGTATTCcTTgTcdTGTGTGG24.2
M07623_2.00CIS-BPGTCTaaTAGAGCTGGGcTGCA23.9
ZNF157 (MA2331.1)JASPARAGGCCAGCAATAtACcTTtAC23.8
M07572_2.00CIS-BPAaTaCAGCCCTAGACTCTCCC23.8
ZN470.H14CORE.0.PSG.APSGAHOCOMOCOGnnhTCTTCTTCTGtTtaCT23.6
M07651_2.00CIS-BPCggaaacCCmcCaGGgCCcgGAGCCtTTt23.3
ZN567.H14CORE.0.P.CPCHOCOMOCOAAcAgtCAAtAgAAacAGACCCagAgATG23
M07649_2.00CIS-BPCccsCCCCccCCbCAGGAATgcc23
M08356_2.00CIS-BPAGAATTAGTGAAcTGGAAGAt23
M07570_2.00CIS-BPAGTTGAGTGACTGcgGTT22.9
ZNF26.H14CORE.0.P.BPBHOCOMOCOATGTCtAGCAtaCAATCAAAAAT22.8
M07689_2.00CIS-BPCCCaGGCtCCTTCCatCTTgTgGCTC22.8
M07705_2.00CIS-BPGCCAAtCACTGAGACAACAaGTaTTGC22.8
ZN799.H14CORE.0.P.CPCHOCOMOCOtgGCAAtACttGTTGTCTCAGTGaTTGGCtT22.6
M07712_2.00CIS-BPacaTTTTttTaATnTAAaaAaAbAttTrCa22.6
ZNF140 (MA1589.2)JASPARAGGAGtGGAATTGCTGGgt22.5
ZNF582 (MA1983.2)JASPARTcTgTTACTTGCAGCCaAA22.5
ZN316.H14CORE.0.P.CPCHOCOMOCOCCTGCCCTCAgGGAGCTTaCAnTC22.5
ZN789.H14CORE.0.P.CPCHOCOMOCOTCCCACTTCTGACACCa22.5
M07621_2.00CIS-BPCAGCCGCCCCGTCCG22.5
M07678_2.00CIS-BPccngccyCcTCcTCCCcCTtCCCaCcC22.5
ZN285.H14CORE.0.P.CPCHOCOMOCOGCCaATTtCcgTGGTGTAAATAC22.3
M07629_2.00CIS-BPatTTTGCCTTGCTGGga22.3
ZN879.H14CORE.0.P.CPCHOCOMOCOTTtGGtgCATaTATgTTtAntaTTgTTAT22.2
ZN33B.H14CORE.0.P.CPCHOCOMOCOgAAGGaATTtAATabAGGGaaTTgg22.1
ZN680.H14CORE.0.P.BPBHOCOMOCOccatGhCCAAGAAGAATgAGG22.1
ZNF23.H14CORE.0.P.CPCHOCOMOCOCCaTGGCAGACATcACTAATCaAT22.1
ZN347.H14CORE.0.PSGI.APSGIAHOCOMOCOAAgGGGATGCGAhaGGGAcg22
ZNF528 (MA1597.1)JASPARCcCAGGGAAGCCATTTC21.9
ZN493.H14CORE.0.PSGI.APSGIAHOCOMOCOcnagnCAtCCaGAaCAGTGCCCa21.9
M07760_2.00CIS-BPGCTGGGAGGCTA21.9
ZN746.H14CORE.0.PSG.APSGAHOCOMOCOcCCCCCCCCccccgCCCTCCCcCC21.8
ZN836.H14CORE.0.PSG.APSGAHOCOMOCOaaTCAGCCACGCCAC21.7
ZNF12.H14CORE.1.M.CMCHOCOMOCOGCTATAACGAGCAtcCCC21.7
ZN724.H14CORE.0.PSG.APSGAHOCOMOCOCGaCgTgCTTGGACTT21.6
M07599_2.00CIS-BPcTTcTTGAGGaCCTcGATgA21.6
M07594_2.00CIS-BPCAgaCccCCaaAcaCcgCaCaCacbaCAt21.5
M04615_2.00CIS-BPTcGaTCGaavggnvcAGTCAT21.3
ZN519.H14CORE.0.P.CPCHOCOMOCOGCCGCcGCcGCcGCCGCCC21.2
ZN880.H14CORE.0.P.CPCHOCOMOCOttgCTGCACTCTCCCtCtCCCA21.2
ZFP28.H14CORE.0.P.BPBHOCOMOCOTCTATTTCTTCTtGwGTCAgTTT21.1
ZN765.H14CORE.0.P.CPCHOCOMOCOGGcCCTGGGCCTaGCCCAgGAAA21
M08305_2.00CIS-BPTCCTCCAGGAAGCCc21
M04527_2.00CIS-BPCaCCCATGGCCGCGaCc20.9
ZN551.H14CORE.0.PSG.APSGAHOCOMOCOGAGCctCCAAtAAAAACGcTG20.8
M07659_2.00CIS-BPGTCTtCcaAGtaGCTGGtGTT20.8
M07773_2.00CIS-BPTTtCtCccCCtccCCctCcCCnTycCTcTc20.8
M07620_2.00CIS-BPTtGgATcTGTgGaTtcATgtCTTTCAT20.7
ZNF282 (MA1154.2)JASPARCTTTCCCacAACACG20.6
ZN274.H14CORE.1.P.BPBHOCOMOCOGAGAAcTCAtACTGGaGAGAA20.6
ZN485.H14CORE.0.P.CPCHOCOMOCOTCaGsvTCCTCCCAhATaTCCCC20.6
ZNF707 (MA1715.1)JASPARCCCCACTCCTGGTAC20.5
ZN570.H14CORE.0.P.CPCHOCOMOCOAGAtATAAtaCAAGAAAAc20.5
ZN211.H14CORE.1.P.CPCHOCOMOCOttTGTtAatATGGTATATAt20.4
ZN669.H14CORE.0.P.BPBHOCOMOCOGhAGCtAGAGcGGTCATCgcCCA20.4
ZN569.H14CORE.0.SG.ASGAHOCOMOCOGGACTAGTAgTAntantaaAaA20.3
M07593_2.00CIS-BPAGaTGAaGTCAcccbTCTnaC20.3
ZN226.H14CORE.0.PSGI.APSGIAHOCOMOCOaTdaGATAAATaGGcAahCG20.2
ZN500.H14CORE.0.PSGI.APSGIAHOCOMOCOGCGTCTACACtancC20.2
ZN566.H14CORE.0.P.CPCHOCOMOCOAGCTGCAAGGGAGGCtGGgAa20.1
ZNF684 (MA1600.2)JASPARACAGTCCACCCCTT20
ZN132.H14CORE.0.P.CPCHOCOMOCOCctCCtTCCTTCCCtCTcTCCtt20
ZN726.H14CORE.0.PSGI.APSGIAHOCOMOCOGccsTAGgGATTGCTTa20
ZN823.H14CORE.0.P.CPCHOCOMOCOGAcAgaGGgAttGcAanAGAGAA20
ZN107.H14CORE.0.PSGI.APSGIAHOCOMOCOaCTTTGTGCACTTGCgC19.9
ZN229.H14CORE.0.SG.ASGAHOCOMOCOACCTTCAGtaAAG19.9
ZN682.H14CORE.0.P.BPBHOCOMOCOaGAACCCTCCcgGGCTAAGCCCC19.9
ZN160.H14CORE.0.SG.ASGAHOCOMOCOGAAaTGAAAAATgCAaTaGA19.8
M07605_2.00CIS-BPthtTttttTthaTtTTTTttTTTtTTTTTT19.8
M07765_2.00CIS-BPCCTTTTTgtTGcTCTTtT19.8
ZN214.H14CORE.0.P.BPBHOCOMOCOgncaTTtTTGAGGacnTTGATGA19.7
M07576_2.00CIS-BPTGCTGCCCAATTCAT19.7
ZN814.H14CORE.0.PSG.APSGAHOCOMOCOTTAGGGAGGGA19.6
ZN595.H14CORE.0.P.CPCHOCOMOCOATGttAgAGgtTTaAAaCACTtg19.5
M07601_2.00CIS-BPCAGATGCCgGCaccaTGCTTC19.5
M08281_2.00CIS-BPCCTCtgGGCCTG19.5
M07734_2.00CIS-BPGvaGCnGCAGCnGCtGccGcg19.4
ZNF343 (MA1711.2)JASPARCGCTTCaCCdCGGCa19.3
ZN732.H14CORE.0.PSG.APSGAHOCOMOCOgCGCCAGTGCCaaGGgg19.3
ZN140.H14CORE.1.SM.BSMBHOCOMOCOGCAATTCCGCTC19.2
ZN492.H14CORE.0.P.CPCHOCOMOCOGAGCAAcAAAAAGGcA19.2
ZN75D.H14CORE.0.P.BPBHOCOMOCOGTGGGAAAbCCtcCc19.2
M07706_2.00CIS-BPgbcttTGAGangGcCcagaTGTTGGacTT19.2
M07778_2.00CIS-BPTttmcttnggkgccActtGAcTCcCTggGcTwaagt19.2
ZN382.H14CORE.0.P.BPBHOCOMOCOTGTaGGgTCTGTAGTgaTGtCtC19.1
M07632_2.00CIS-BPCAGTATGAGTTCTCt19.1
M07775_2.00CIS-BPGCCGAAACCCGG19.1
ZN506.H14CORE.0.P.CPCHOCOMOCOTtTGGGGGCTCatcCngG18.9
ZN140.H14CORE.0.P.BPBHOCOMOCOccAGCAATTCCgCTCC18.8
ZN483.H14CORE.0.P.CPCHOCOMOCOcAGgGAGCTGCAGbnnC18.7
ZN549.H14CORE.0.P.BPBHOCOMOCOatTCATGAAttGGGCAGC18.7
ZN707.H14CORE.0.P.BPBHOCOMOCOCACCCCACTCCTGGT18.7
ZN320.H14CORE.0.P.BPBHOCOMOCOGtGGGrcCaGGGGGCCagtG18.6
ZN684.H14CORE.0.P.CPCHOCOMOCOACAGTCCACCCCTT18.6
M07622_2.00CIS-BPaTTTAGwCCATTTAC18.6
M07627_2.00CIS-BPCAGTCACCtTCtGG18.6
M07631_2.00CIS-BPTTTCAAAAhTGCTaTGG18.6
M07772_2.00CIS-BPTGCCCTAGCcktggcTtt18.6
M07692_2.00CIS-BPCATCTsaggGTCTgcTTbTattgabtgtT18.5
ZN841.H14CORE.0.PSGI.APSGIAHOCOMOCOGGTGaagTGGvTGCTgCT18.4
M08378_2.00CIS-BPTtTCtaTTTCtTCtgdtGT18.4
ZN606.H14CORE.0.PSG.APSGAHOCOMOCOgaGGAAACCGCgvC18.3
ZN776.H14CORE.0.P.CPCHOCOMOCOAGCAGATGCCgGCaccaTGCTTC18.3
M07586_2.00CIS-BPGGtGGGgGCGTCCCCC18.3
M07656_2.00CIS-BPTGTCTAGTAaGTCt18.3
M07761_2.00CIS-BPCTGATTATTCACCCA18.3
ZN418.H14CORE.0.P.BPBHOCOMOCOTGCTTTTaGCcTCTg18.2
ZN484.H14CORE.0.P.CPCHOCOMOCOAggGGAnAGAgGAACAdGttAAA18.2
ZNF14.H14CORE.0.P.CPCHOCOMOCOTCTgACCAGTAaACCCAGGCA18.2
ZNF41.H14CORE.0.P.CPCHOCOMOCOCAAGGgAataagvCACCATGAGbgA18.2
M07583_2.00CIS-BPGAATTTaTCACCACT18.2
M07638_2.00CIS-BPTGCAGCTCCCTg18.2
ZNF320 (MA1976.2)JASPARGgGGGgcCcgGGGGCctgTG18.1
ZFP82.H14CORE.0.P.CPCHOCOMOCOcTTCCAkTTcActAAtTcTCtCTTC18.1
ZN180.H14CORE.0.P.CPCHOCOMOCOaGGGagkGGtGGGGaCTGbGGCraa18.1
ZFP69.H14CORE.0.P.CPCHOCOMOCOGTgGCTGGAaACAAGaTAA18
M07578_2.00CIS-BPCAACTCTCC18
M08267_2.00CIS-BPAGTTTCrtTTTCc18
ZNF135 (MA1587.1)JASPARCCTCGACCTCCtga17.9
ZNF136 (MA1588.1)JASPARgaATTCTTGGTTGaC17.9
ZN133.H14CORE.0.P.BPBHOCOMOCOgaaGgtCCAACAtAtGTATAATTGG17.9
ZN197.H14CORE.0.P.CPCHOCOMOCOtctTctctTTGTCTTTgGTgtTc17.9
ZN273.H14CORE.0.P.CPCHOCOMOCOGAGCTaCCCaCTCCgGGgCtcC17.9
ZN587.H14CORE.0.P.CPCHOCOMOCOTGGCGCCCAACaTgGGgC17.9
M07636_2.00CIS-BPCTgCCCTGGGAyTTT17.9
ZN101.H14CORE.0.P.CPCHOCOMOCOttnTnnttGAcACaGGGGCcaC17.8
ZN234.H14CORE.0.PSG.APSGAHOCOMOCOGATAAATaGGaGAtC17.8
ZN713.H14CORE.0.SM.BSMBHOCOMOCOaGACGACTGCCACGAAA17.8
ZN813.H14CORE.0.SG.ASGAHOCOMOCOgtGAAGGAgGGrGg17.8
M07767_2.00CIS-BPAAAAaCCCCAGA17.8
M08380_2.00CIS-BPTgCCTTTTTbTTGTCcTT17.8
M07617_2.00CIS-BPGAAATGaCTTCCCt17.7
ZNF418 (MA1980.1)JASPARrGAGGCTAAAAGCA17.6
M04531_2.00CIS-BPGCCGCCATtTTG17.6
M07657_2.00CIS-BPCCctgCATGTCCCC17.6
M07730_2.00CIS-BPcctcaGgTTgTaGGaatGctGag17.6
M07764_2.00CIS-BPCTGCaCTCTCCCcttCCChccchnc17.6
ZN416.H14CORE.0.P.CPCHOCOMOCOTGACTGGGCcAcgGGgTGCCCAG17.5
ZN816.H14CORE.0.P.BPBHOCOMOCOAaraagGGGGACATGcngGG17.5
M07697_2.00CIS-BPTTAGAgATTcaTTTCTTT17.5
ZN141.H14CORE.0.P.CPCHOCOMOCOGGGGGACGCcCCCaCC17.4
ZN454.H14CORE.0.P.BPBHOCOMOCOCCaTCtGGgCCTGGtgCCtTTT17.4
ZN626.H14CORE.0.P.CPCHOCOMOCOaTAhGGTTcTTgTGgtCCCTG17.4
ZN782.H14CORE.0.P.CPCHOCOMOCOcttTgAGggGGCCCAGATGTTGG17.4
M07710_2.00CIS-BPATTaAATTCCcTctntTtaAA17.4
ZN605.H14CORE.0.P.CPCHOCOMOCOcCAGAtctTGGTtTCTAAataCC17.3
ZN610.H14CORE.1.P.CPCHOCOMOCOtttaCaGCCGCTCCCCaTygCT17.3
ZN677.H14CORE.0.P.CPCHOCOMOCOgCTGTTCTTaTcggGgTtC17.3
M07635_2.00CIS-BPgTTTTCTTGtattaTaTcT17.3
M07650_2.00CIS-BPgtcTATAAAaggggggccgcggsg17.3
M07668_2.00CIS-BPCbGCCCCTttCCtTtC17.3
M07754_2.00CIS-BPCgCCCCCTCCC17.3
ZN440.H14CORE.0.P.CPCHOCOMOCOGncaaAAGCAGAACmACTgaT17.2
M07661_2.00CIS-BPACCCCACTCCTGGT17.2
M07748_2.00CIS-BPGGGCACCCCnCcngCCC17.2
M08382_2.00CIS-BPCTGTTCTTdTcgGggTcC17.2
ZN669.H14CORE.1.P.CPCHOCOMOCOcGGTCATCGcCC17.1
ZN701.H14CORE.0.P.BPBHOCOMOCOGAGcAccaAaGGGGGgA17.1
M07716_2.00CIS-BPCTCccTCCCTCCCt17.1
ZIM3.H14CORE.0.P.BPBHOCOMOCOgGgTTTCTGTTgct17
ZN274.H14CORE.0.SM.BSMBHOCOMOCOGAGAACTCAtA17
ZN468.H14CORE.0.P.CPCHOCOMOCOGGGAGGGGGcG17
M07702_2.00CIS-BPCTTCTTCCTTgg17
ZNF675 (MA1714.2)JASPARGGvhtaGGAGGaCAAAATG16.9
ZN689.H14CORE.0.PSG.APSGAHOCOMOCOaGCCaAtGCAATAaggCAAGa16.9
ZN708.H14CORE.0.P.BPBHOCOMOCOAagbAGGtACAGCt16.9
M07615_2.00CIS-BPACCCCCCCA16.9
M07655_2.00CIS-BPCCCCTCGTGG16.9
M07686_2.00CIS-BPTagCTTGTTTCCAGcCaC16.9
ZNF547 (MA2334.1)JASPARTGCTAATGCAaCA16.8
ZN251.H14CORE.0.PSGI.APSGIAHOCOMOCOTTgCACTTAaCaCcbTntC16.8
ZN615.H14CORE.0.P.CPCHOCOMOCOAAcaACtTTATCAaCtAGaG16.8
ZN681.H14CORE.0.P.CPCHOCOMOCOcccttctgtgTCCtCvTCCTT16.8
M04503_2.00CIS-BPGcTTTTCCCAC16.8
M07596_2.00CIS-BPTgTACTGCtkdCCcTvCCAgGaC16.8
M07614_2.00CIS-BPCCcCagbCCCCdCCccbCCC16.8
M07565_2.00CIS-BPTGGGGGCTCvtcC16.7
M07667_2.00CIS-BPCacwgGCCCCcgGgcCCC16.7
M07687_2.00CIS-BPaTaaGGtTcttgTGccCCCTGC16.7
M07739_2.00CIS-BPTgTccATGAACATkGTatac16.7
AC0232:ZNF33B:C2H2_ZFVierstraATTaAATTCCcTctntTtaA16.7
ZNF530 (MA1981.2)JASPARGaAaGGavAGGGGC16.6
ZN283.H14CORE.0.P.CPCHOCOMOCOGctnGGCTGATCTgGgcTGGcc16.6
ZN749.H14CORE.0.P.CPCHOCOMOCOcvcCnCtgACCCCAaCcTtcC16.6
ZN793.H14CORE.0.P.BPBHOCOMOCOCTTGGGGCTCtgkgggcttc16.6
M07642_2.00CIS-BPCcGCcGCcGCCGCCC16.6
M07654_2.00CIS-BPtGCCcCCTkGTGGC16.6
M07728_2.00CIS-BPTCtnaacTGGTgccACCA16.6
ZNF274 (MA1592.2)JASPARTaTGAGTTCTCg16.5
ZNF75A (MA2097.1)JASPARGcTTTTCCCAC16.5
ZN433.H14CORE.0.P.CPCHOCOMOCOagTTActACtGcgGTCmT16.5
ZN534.H14CORE.0.P.CPCHOCOMOCOGgGGtTCTtGCCCCcC16.5
ZN547.H14CORE.0.P.BPBHOCOMOCOTGCtAATgCwGCAgGCAnaC16.5
ZN728.H14CORE.0.PSG.APSGAHOCOMOCOGTtGGwdtsaCTGgaaCCAGC16.5
M07566_2.00CIS-BPTCvAACCATCCTTGC16.5
M07580_2.00CIS-BPCCAaTTaTACnTntgTTGgacC16.5
M07591_2.00CIS-BPaGvTTTCTGTT16.5
M07598_2.00CIS-BPGAcAatavCAAGTGtTGaC16.5
M07683_2.00CIS-BPGTGGGAAAgCCT16.5
ZFP14.H14CORE.0.P.CPCHOCOMOCOGCvTTCCaGtgcCTCC16.4
ZN573.H14CORE.0.P.CPCHOCOMOCOTCAGcCadGGcACAchcAgGa16.4
ZN665.H14CORE.0.PSG.APSGAHOCOMOCOGAaACctcCcgCGACGcc16.4
ZN671.H14CORE.0.P.CPCHOCOMOCOgaatgGAgTGGAwtGGartGggg16.4
M07587_2.00CIS-BPCCctCCCCagTCgaGcC16.4
M07685_2.00CIS-BPTTTCCAGCCAC16.4
M07763_2.00CIS-BPAGAGGTTTCTT16.4
ZNF701 (MA1987.2)JASPARGAGCAccaaaGGGGGgA16.3
ZN432.H14CORE.0.P.CPCHOCOMOCOTTAAACAaTTGCCdcTgATTGTT16.3
ZN577.H14CORE.0.P.BPBHOCOMOCOCarvgCCcCTTGAGgcCt16.3
ZN878.H14CORE.0.PS.APSAHOCOMOCOCanagCAGGACAAAAA16.3
M07658_2.00CIS-BPCCGCCGCvGCCgCCg16.3
M07677_2.00CIS-BPcCcGCcTCCctTsCcGC16.3
M07770_2.00CIS-BPCTTGGACTT16.3
ZF69B.H14CORE.0.P.CPCHOCOMOCOGTGGCTGGAaA16.2
M07673_2.00CIS-BPgTCCCCcgaGTCaTC16.2
M07726_2.00CIS-BPACtAcTGTggTaATT16.2
ZNF816 (MA1719.2)JASPARtGGGGACATGcngGG16.1
Z324A.H14CORE.0.P.BPBHOCOMOCOaTcaAACCATCCTTkgCTgsc16.1
ZN211.H14CORE.0.P.BPBHOCOMOCOaTGGTATATAt16.1
ZN479.H14CORE.0.P.CPCHOCOMOCOGGAtGACtcgGgGGat16.1
ZN845.H14CORE.0.PSG.APSGAHOCOMOCOGGGaCTgGAGAGcadgtggGg16.1
M07633_2.00CIS-BPTcbttgTGCctTTGCCaT16.1
M07722_2.00CIS-BPCaTTtTGtCCTCCtgnbCC16.1
ZNF75D (MA1601.2)JASPARGTGGGAAAgCCT16
ZN429.H14CORE.0.P.CPCHOCOMOCOGCCCATTgbgnCCTcTGCCT16
ZN530.H14CORE.0.P.BPBHOCOMOCOCnGcCCCTbkCCtTgCc16
ZN611.H14CORE.0.P.CPCHOCOMOCOGGAGcGCCtatgaacggccG16
ZNF79.H14CORE.0.P.CPCHOCOMOCOAAAgAAAtkAATcTCTAA16
M07588_2.00CIS-BPCCTTCCChCtyTCC16
M07663_2.00CIS-BPGAcACaGGGGCcAC16
M07672_2.00CIS-BPCTTTAGCTGcAT16
M07693_2.00CIS-BPCcTgnGTGTGCChtggCTg16
M07741_2.00CIS-BPAcATaTCCCCATTCC16
ZNF324 (MA1977.2)JASPARAGhcAAGGATGGTT15.9
ZNF669 (MA1985.1)JASPARgGGgCGAtGACCacT15.9
ZN548.H14CORE.0.P.BPBHOCOMOCOCaAGatcaaGGtGgCdGCAGGG15.9
M07579_2.00CIS-BPCATCTCCAGga15.9
M07639_2.00CIS-BPTGGCGCCCAacrtgG15.9
M08318_2.00CIS-BPcccCAGCTaCTCC15.9
ZIK1.H14CORE.0.P.CPCHOCOMOCOAaAAtGGCAAAaGCAaaavgA15.8
ZN266.H14CORE.0.P.BPBHOCOMOCOTaCTCACAGgtCCc15.8
M07691_2.00CIS-BPCCTGChGcCaCCbtGvcCttG15.8
M07715_2.00CIS-BPGCCGCCGCCdTgG15.8
M08400_2.00CIS-BPAACCTCTAACAT15.8
AC0318:ZNF33A:C2H2_ZFVierstraTCAGTGCAC15.8
ZN850.H14CORE.0.PSG.APSGAHOCOMOCOAaaCCtGAGAGGGGCTtC15.7
ZN208.H14CORE.0.SGI.ASGIAHOCOMOCOACgAACCcTAAt15.6
M07608_2.00CIS-BPtdgggtCTGTAcTgnTgtCTC15.6
M07634_2.00CIS-BPAAACGTCCACg15.6
ZN267.H14CORE.0.P.CPCHOCOMOCOctcCctccsGGCaTGTGaaGAdGggCctG15.5
ZN699.H14CORE.0.PSG.APSGAHOCOMOCOcvGGcgatGAAGGGTG15.5
M07624_2.00CIS-BPGccGGCATCTGGtg15.5
M08284_2.00CIS-BPCGCCGCCtGmgTTC15.5
M08342_2.00CIS-BPGCTCtGbCTCcctC15.5
ZNF680 (MA1729.2)JASPARCCAAGAAGAAT15.4
ZN568.H14CORE.0.SG.ASGAHOCOMOCOAAGAACAGC15.4
M07619_2.00CIS-BPggCCvGGCCaGAtCAGCCnagcc15.4
M07746_2.00CIS-BPannTgGCGCCCAACATkGgbc15.4
M07774_2.00CIS-BPCCCTACCTC15.4
ZN510.H14CORE.0.PSG.APSGAHOCOMOCOTGATGAGGaAaCtantArGabGt15.3
ZN891.H14CORE.0.P.CPCHOCOMOCOGGGCTAGAAGcC15.3
M07585_2.00CIS-BPCTGCaGAGCCCt15.3
M07690_2.00CIS-BPgGGGCGAcGACCacT15.3
AC0542:ZNF75A/ZNF75D:C2H2_ZFVierstraGTGGGAAAAgC15.3
ZN189.H14CORE.1.P.BPBHOCOMOCOctttgCmchtgCTGTTCCctc15.2
ZN528.H14CORE.0.P.BPBHOCOMOCOAGGGAAGcCATTTCT15.2
ZN709.H14CORE.0.P.CPCHOCOMOCOGAGtCcCAGAAGGaGadGAG15.2
M07600_2.00CIS-BPGCATTAGC15.2
M07759_2.00CIS-BPGGAGCGCCycnGccsggC15.2
AC0302:ZIM:C2H2_ZFVierstraGgTTTCTGTT15.2
ZIM3 (MA1709.2)JASPARAACAGAAAcC15.1
ZNF85 (MA1720.2)JASPARgAGATTACagCA15.1
ZN678.H14CORE.0.PSG.APSGAHOCOMOCOGcAatGGGtAGGAcAGCA15.1
ZN766.H14CORE.0.P.BPBHOCOMOCOCcaAAgAAACCTCtTtT15.1
ZNF20.H14CORE.0.PSG.APSGAHOCOMOCOAGTACTCTCCT15.1
M07577_2.00CIS-BPATGGTATAT15.1
M07592_2.00CIS-BPCvTTCCnGkgCCTCC15.1
M07731_2.00CIS-BPTCcchgCaGCAacAnC15.1
M07744_2.00CIS-BPGCACAGCgcCC15.1
AC0192:ZIK:C2H2_ZFVierstraTcbttgTGCctTTGCC15.1
ZFP14 (MA1972.1)JASPARGGAGGcmCnGGAanG15
ZN561.H14CORE.0.P.CPCHOCOMOCOCtGCAGAAAGGGtgCnccc15
ZN584.H14CORE.1.P.CPCHOCOMOCOATTTCAAAAhTGC15
ZN721.H14CORE.0.PSG.APSGAHOCOMOCOAgAAAAAAGCtgccCC15
ZN793.H14CORE.1.P.CPCHOCOMOCOCTTGGGGCTC15
ZNF12.H14CORE.0.P.CPCHOCOMOCOGCTGCtGTAACAAAharcCCc15
M07735_2.00CIS-BPGTAGCTCCT15
ZNF454 (MA1712.2)JASPARGGCTCcsGGcCCcggkG14.9
ZN610.H14CORE.0.P.BPBHOCOMOCOCmGCCGCTCCcC14.9
ZN616.H14CORE.0.P.CPCHOCOMOCOGhgnrGGTGAGCgtGagtagtCc14.9
ZNF57.H14CORE.0.PSG.APSGAHOCOMOCOaGGgvGGGGAGGGTGTa14.9
Z585A.H14CORE.0.P.CPCHOCOMOCOaaaaaaaAAaAGAgAACAGA14.8
ZN157.H14CORE.0.P.BPBHOCOMOCOChttGtChTGGbaGGghhagCAgTata14.8
M07665_2.00CIS-BPGCTGTACCT14.8
M07753_2.00CIS-BPGcdTtcCTTGGCTTGTG14.8
M08372_2.00CIS-BPCAATTCcGCTC14.8
Z705G.H14CORE.0.P.CPCHOCOMOCOattaGAgGtTtaTTycatTc14.7
ZFP90.H14CORE.0.P.CPCHOCOMOCOACTGCTTTaGCTG14.7
M07637_2.00CIS-BPCthakTCTTCTTgGnC14.7
M07676_2.00CIS-BPCctCtgCaCabGCCgkcaG14.7
M07725_2.00CIS-BPAAGcACTGTACT14.7
M07751_2.00CIS-BPGgtTCTCgCCCCcC14.7
ZN714.H14CORE.0.P.CPCHOCOMOCOgcrgTGCTGCT14.6
ZNF85.H14CORE.1.P.BPBHOCOMOCOaGAAaacTGaaGtAATCT14.6
M07647_2.00CIS-BPTCGACCTCC14.6
ZNF682 (MA1599.2)JASPARGGCcAAGCCCC14.5
ZNF211 (MA1974.2)JASPARtATATACCAt14.5
Z587B.H14CORE.0.SG.ASGAHOCOMOCOGGACAcTGgc14.5
ZN100.H14CORE.1.P.BPBHOCOMOCOCcAhGGCGGCGGC14.5
ZN394.H14CORE.0.P.CPCHOCOMOCOGGAGwAGCTG14.5
ZN431.H14CORE.0.P.CPCHOCOMOCOGGGCTaGAAGcC14.5
ZN527.H14CORE.0.P.CPCHOCOMOCOTttCTGTtgCTtgCaaCC14.5
ZNF66.H14CORE.0.PG.APGAHOCOMOCOCcgCTGCAtACAcc14.5
ZNF83.H14CORE.0.G.CGCHOCOMOCOCTTtTTcTCTTdc14.5
M07613_2.00CIS-BPTCanTcAgTCAdTCA14.5
M07688_2.00CIS-BPGAGCCCCAAGc14.5
M07641_2.00CIS-BPACTCACAGgcCC14.4
M07714_2.00CIS-BPgCCCATTgnbdCCtaTgCC14.4
M07736_2.00CIS-BPtTtCCTGTTChtdcnnyTtyC14.4
ZFP37.H14CORE.0.P.CPCHOCOMOCOAGAAATGcttggahcaataA14.3
ZN417.H14CORE.0.P.CPCHOCOMOCOCAntTGGCGCC14.3
ZN662.H14CORE.0.P.CPCHOCOMOCOatCTAGCCCctgCCTaCC14.3
ZN729.H14CORE.0.SG.ASGAHOCOMOCOTGCGGCAg14.3
M07597_2.00CIS-BPcCTTTTGTTTTT14.3
M07616_2.00CIS-BPGCCGCTCC14.3
M07666_2.00CIS-BPCTAgCCCcbgCCTaCC14.3
M07701_2.00CIS-BPCTgGGCtanGCcCtGsC14.3
M07718_2.00CIS-BPcCchcscnGGccAAGCCC14.3
ZN613.H14CORE.0.P.CPCHOCOMOCOCtntttTTTTTTTttTTtbt14.2
ZN773.H14CORE.0.PG.APGAHOCOMOCOCaCyAmmnaCtCCaCCttcAc14.2
M07723_2.00CIS-BPGCTCCCTGC14.2
M08371_2.00CIS-BPCAGaCTTgTCcTc14.2
ZN337.H14CORE.0.P.CPCHOCOMOCOGaaTTtgTCaccaCTaGACCaGC14.1
ZN543.H14CORE.0.P.CPCHOCOMOCOCCcbGCCnccbGGCCCCCTCC14.1
ZN860.H14CORE.0.P.CPCHOCOMOCOGCTCCCTGC14.1
M07571_2.00CIS-BPATGCnyCTTTCTA14.1
M07752_2.00CIS-BPATTACTAATgAAGtT14.1
AC0033:ZNF354A/ZNF354B:C2H2_ZFVierstraTTTAgtCCATTTA14.1
ZNF708 (MA1730.2)JASPARGCTGTgCCT14
ZN223.H14CORE.0.P.CPCHOCOMOCOGGtGGtgCAGGCa14
ZN480.H14CORE.0.P.CPCHOCOMOCOCCaaTChChvnhnCaCatTCc14
ZN75A.H14CORE.0.SM.BSMBHOCOMOCOGTGGGAAAAgc14
M07606_2.00CIS-BPGCagTGCTGCT14
M07612_2.00CIS-BPCCgCcTCC14
M07711_2.00CIS-BPGgCTTCtAGCCC14
M07762_2.00CIS-BPGGCTTCtAGCC14
ZN816.H14CORE.1.P.CPCHOCOMOCOAaraAgGGGGACATG13.9
ZNF7.H14CORE.0.P.BPBHOCOMOCOctTGcCarCACTTgntatTgtC13.9
M07732_2.00CIS-BPTGCTGCAC13.9
M07750_2.00CIS-BPCttCTGACACCa13.9
ZNF677 (MA2101.1)JASPARgAtAAGAACAG13.8
ZN257.H14CORE.0.P.BPBHOCOMOCOGctCTTGCCTC13.8
ZN454.H14CORE.1.SM.BSMBHOCOMOCOCGCCGGCGCCa13.8
ZNF44.H14CORE.0.P.CPCHOCOMOCOGbTaTTGCwGtdgaGAA13.8
M07652_2.00CIS-BPGGcCCCCTCC13.8
M07717_2.00CIS-BPgCTCCGCCcC13.8
ZN124.H14CORE.0.P.CPCHOCOMOCOCCTTCTTCCTTg13.7
ZN264.H14CORE.0.P.BPBHOCOMOCOAAGGGCACTAATCCcAt13.7
ZNF28.H14CORE.0.P.CPCHOCOMOCOggGndggtGGGGTGCC13.7
ZNF77.H14CORE.0.P.CPCHOCOMOCOcTtccCACTtCACCctvthaAa13.7
M07653_2.00CIS-BPaTtTAttCCATT13.7
M07662_2.00CIS-BPtgTCCTCCctngtcC13.7
ZNF610 (MA1713.2)JASPARccGCCGCTCC13.6
ZN676.H14CORE.0.PSG.APSGAHOCOMOCOGCGAcagtaCtcmgCaC13.6
ZNF34.H14CORE.0.P.CPCHOCOMOCOgcmhtCAGaCTTkTCcT13.6
M07573_2.00CIS-BPTGCtGGAATCTc13.6
M07628_2.00CIS-BPagTgGTTCTGCT13.6
M07694_2.00CIS-BPtttCTGTtgCttgCacCC13.6
M07757_2.00CIS-BPCcGCGgCC13.6
ZN343.H14CORE.0.PSM.APSMAHOCOMOCOCGCTTCAC13.5
ZN766.H14CORE.1.P.BPBHOCOMOCOAAGAAACCt13.5
M07581_2.00CIS-BPCctGTgCttCtbTCCcc13.5
M07733_2.00CIS-BPtTTCTCTbTctCc13.5
ZNF257 (MA1710.2)JASPARGAGGCaAGgG13.4
ZNF214 (MA1975.2)JASPARTCATCAAbGtCCT13.4
M08366_2.00CIS-BPGTtTGCTGCTtnC13.4
M08368_2.00CIS-BPTCctbdCyGGCAgCtG13.4
ZN674.H14CORE.0.P.CPCHOCOMOCOtGGcGvTCCcbgG13.3
M07574_2.00CIS-BPCACCCCTTt13.3
M07575_2.00CIS-BPTcCAaGGhGGCcaGC13.3
M07743_2.00CIS-BPTGATGAC13.3
M08245_2.00CIS-BPTGGTGbcttatTcCC13.3
M08341_2.00CIS-BPCaGaGCCAbgTgGggtgC13.3
ZN177.H14CORE.0.SM.BSMBHOCOMOCOTGActdtnccbtTCGATCgA13.2
ZNF3.H14CORE.0.P.BPBHOCOMOCOtGtTTGtTGAnTGAaTvantg13.2
ZN215.H14CORE.0.PSG.APSGAHOCOMOCOCcGCGCCTCC13.1
ZNF43.H14CORE.0.PSGI.APSGIAHOCOMOCOCtAGATTGCTcC13.1
M07630_2.00CIS-BPTTTCTGC13.1
ZNF331 (MA1726.2)JASPARTGCAGAGCC13
ZNF417 (MA1727.2)JASPARGGCGCCA13
ZN189.H14CORE.0.P.BPBHOCOMOCOtgCTGTTCCc13
ZN621.H14CORE.0.P.CPCHOCOMOCOAAAgTCCCAGGGaAG13
ZNF26.H14CORE.1.P.BPBHOCOMOCOAcAATGtCtAGCAt13
M07644_2.00CIS-BPACTcCAC13
M07699_2.00CIS-BPGGTTTaT13
M07703_2.00CIS-BPCCGACGg13
M07724_2.00CIS-BPTTCCTCc13
M07755_2.00CIS-BPtCacGCTCACCcnc13
M07758_2.00CIS-BPcGCCCTC13
ZNF766 (MA2098.1)JASPARAAGAAACCt12.9
ZFP1.H14CORE.0.PSM.APSMAHOCOMOCOTTttATaCCCAGChaaacTatC12.9
ZNF18.H14CORE.0.P.BPBHOCOMOCOGGTGTGAAC12.9
M07684_2.00CIS-BPcctgttcTTGaCTTT12.9
ZNF416 (MA1979.2)JASPARTaTCTGGGCA12.8
ZN383.H14CORE.0.P.CPCHOCOMOCOGcgGAGCvasaAgakGGaaGGrg12.8
ZNF12.H14CORE.2.S.CSCHOCOMOCOTaTAACAAahAggnC12.8
M07618_2.00CIS-BPCCtttGgTGCTC12.8
M07664_2.00CIS-BPaAtnTTnmTATTTCC12.8
M07681_2.00CIS-BPGTGCcTGcTCagG12.8
M07781_2.00CIS-BPacGTACCACAtT12.8
ZNF184 (MA2120.1)JASPARTAGAAAGgnGCAT12.7
ZN708.H14CORE.1.P.BPBHOCOMOCOAGGtACAGC12.7
ZNF30.H14CORE.0.P.CPCHOCOMOCOGGtTGtaGGACTGagGTcC12.7
M07679_2.00CIS-BPTGCGATCtC12.7
M07698_2.00CIS-BPcCbgCGTCCTT12.7
M07704_2.00CIS-BPtGGtbTCTahnnCCCc12.7
AC0346:ZNF705G/ZNF:C2H2_ZFVierstraGGTTTcT12.7
ZN540.H14CORE.0.P.CPCHOCOMOCOAAACataCACaGaAaCa12.6
ZNF8.H14CORE.0.P.CPCHOCOMOCOGGTACGcChabg12.6
ZN790.H14CORE.0.P.CPCHOCOMOCOTaAcwGCTcCCmnttghTgCt12.5
M08396_2.00CIS-BPGCtcTTTTaTgacc12.5
Zfp961 (MA2126.1)JASPARGGGCGCCA12.4
ZN282.H14CORE.0.P.BPBHOCOMOCOGGdGtGGGAG12.4
ZNF85.H14CORE.0.P.BPBHOCOMOCOTGaaGtAATCT12.4
M07589_2.00CIS-BPttCccTTCC12.4
M07646_2.00CIS-BPTTTTTTTT12.4
M07680_2.00CIS-BPAACCCCAAC12.4
M08252_2.00CIS-BPGCTCCAgCacC12.4
M08373_2.00CIS-BPCcChgbCTCttCcTCt12.4
ZNF667 (MA1984.2)JASPARTTAaGAGCTCA12.3
ZN213.H14CORE.0.P.CPCHOCOMOCOGAAatcAGGCgGCGGg12.3
ZN274.H14CORE.2.P.BPBHOCOMOCOCAtaCtGGaGAGAAacCcTAc12.3
ZN582.H14CORE.0.P.CPCHOCOMOCOGGCTGCAAGTA12.3
ZN716.H14CORE.0.P.CPCHOCOMOCOCagGaAAtaknAancTTActa12.3
M07602_2.00CIS-BPgGTGTGAAc12.3
M07611_2.00CIS-BPcntttgTtnCaGCaGC12.3
M07780_2.00CIS-BPAGCGCCcCC12.2
M07569_2.00CIS-BPGCTTCCCc12.1
M07771_2.00CIS-BPtbTGGtGCtCC12.1
ZNF189 (MA1725.2)JASPARtGCTGTTCC12
M07567_2.00CIS-BPcTCCAcTC12
Zfp809 (MA2125.1)JASPARaTCCCAGCC11.9
ZN554.H14CORE.0.P.CPCHOCOMOCOGCtGAGcCAbgtaGggtaC11.9
ZN778.H14CORE.0.P.CPCHOCOMOCOgGaGtCGgtGTCTG11.9
M07738_2.00CIS-BPTTATAT11.9
ZNF213 (MA2121.1)JASPARgCCCAGGCdGc11.8
ZN682.H14CORE.1.P.BPBHOCOMOCOGGccaAGCCC11.8
M07603_2.00CIS-BPcncCGTCagggcCCc11.8
M07721_2.00CIS-BPCTGGGgC11.8
ZNF354A (MA1978.2)JASPARtAAdtaTAAAtGgactAatT11.7
ZN333.H14CORE.0.P.CPCHOCOMOCOgTGGAgccGAG11.7
ZN788.H14CORE.0.S.CSCHOCOMOCOGTGAAC11.7
ZN350.H14CORE.0.P.BPBHOCOMOCOgcccTTTTATgACC11.6
ZN701.H14CORE.1.P.BPBHOCOMOCOAaGGGGGGAA11.6
ZN205.H14CORE.0.P.CPCHOCOMOCOGAGcgCcaActayctscnaGG11.5
M07745_2.00CIS-BPCaCATTCct11.5
ZN224.H14CORE.0.P.BPBHOCOMOCOAgCccAGgGAgTCaagtggchcC11.4
ZN471.H14CORE.0.PSG.APSGAHOCOMOCOAncTCnaTcTCdtt11.4
ZN614.H14CORE.0.P.CPCHOCOMOCOCTTgATCTaATtgA11.4
ZN749.H14CORE.1.P.CPCHOCOMOCOaCCCCAAcC11.3
M07713_2.00CIS-BPTCTGTacTC11.3
AC0415:ZNF585A:C2H2_ZFVierstraTCTGTacTC11.3
Z780B.H14CORE.0.PSGI.APSGIAHOCOMOCOCTGCTGaa11.2
ZN563.H14CORE.0.P.CPCHOCOMOCOGgnTCCTbdctgGCAgCtG11.2
ZN684.H14CORE.1.M.CMCHOCOMOCOCcntCCaCCCCt11.2
M07595_2.00CIS-BPGcctbtTcCTCtC11.2
M07607_2.00CIS-BPGGCTCCAC11.2
ZN667.H14CORE.0.P.CPCHOCOMOCOGtggCCtTaagAGCTCabC11.1
M07582_2.00CIS-BPTCCCTG11.1
M07669_2.00CIS-BPGGGaaGCtA11.1
M07719_2.00CIS-BPGAGGCAA11.1
ZNF549 (MA1728.2)JASPARTGCTGCCC11
ZNF175 (MA2332.1)JASPARACAGGAAGT11
RBAK.H14CORE.0.P.CPCHOCOMOCOGagAgaAacaggCC11
ZN557.H14CORE.0.P.CPCHOCOMOCOAcCTGgragtGgaTAbTkgtrgG11
M07604_2.00CIS-BPGCTGAcATc11
M07707_2.00CIS-BPTCCCGc11
ZN730.H14CORE.0.P.CPCHOCOMOCOGccaTAGCatCCC10.9
M07626_2.00CIS-BPCTCCCc10.9
ZNF41.H14CORE.1.P.CPCHOCOMOCOCAcAAGGgAg10.8
M07645_2.00CIS-BPCTCCtT10.8
ZN441.H14CORE.0.P.CPCHOCOMOCOGgGCGGAGt10.7
ZN888.H14CORE.0.P.BPBHOCOMOCOAgGGGGGA10.7
ZN250.H14CORE.1.M.CMCHOCOMOCOAGGCCTA10.6
ZN700.H14CORE.0.PSG.APSGAHOCOMOCOaACGGTACt10.5
ZN320.H14CORE.1.P.BPBHOCOMOCOgGGGGCCagt10.4
M08264_2.00CIS-BPTGCAaTTTTt10.4
ZN716.H14CORE.1.P.CPCHOCOMOCOCAGGAAAtA10.1
ZN667.H14CORE.1.P.CPCHOCOMOCOTAAaAGCTCA9.8
ZN790.H14CORE.1.P.CPCHOCOMOCOAcaGCTCCC9.7
M08279_2.00CIS-BPCtccCTCC9.7
AC0154:ZNF354A/ZNF:C2H2_ZFVierstraTAaaaatAAAwgahatAaaT9.7
ZN480.H14CORE.1.P.CPCHOCOMOCOCcCAtTCCc9.6
ZN563.H14CORE.1.P.CPCHOCOMOCOcaGGCAGCag9.5
ZN222.H14CORE.0.P.CPCHOCOMOCOTsTCAGCC9.4
ZN425.H14CORE.0.P.CPCHOCOMOCOGAGGGCc9.3
ZNF354C (MA0130.1)JASPARaTCCAC9
M02666_2.00CIS-BPaTCCAC9
M07695_2.00CIS-BPGTGbCT8.7
AC0412:ZNF33A:C2H2_ZFVierstraGTGbCT8.7
ZN554.H14CORE.1.P.CPCHOCOMOCOGCtGAGcCA7.8
M00250_2.00CIS-BPGcTA3.9

HOCOMOCO tags read as assay + quality grade: P ChIP-seq, S HT-SELEX, M methyl-HT-SELEX, G GHT-SELEX, I SMiLE-seq, B PBM; grade A means two or more assay types agreed. A P-only matrix is a record of where the protein was crosslinked, which need not be a sequence it binds itself.

The factors, elsewhere

how this page was made Drafted by a language model reading the primary literature, then checked: every cited PMID is resolved against PubMed and its citation text regenerated from PubMed's own record, and every claim about the motifs themselves is checked against the matrices in the four source databases. Methods and limits →

Other C2H2 zinc finger families